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Eocene mammals of Africa

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Basilosaurus
Basilosaurus (meaning "king lizard") is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the southeastern United States. The generic name, meaning "king lizard", was given due to the initial misconception about the fossil material as that of a giant reptile. The animal was later found to be an early marine mammal, prompting attempts at renaming the c
Moeritherium
Moeritherium ("Lake Moeris' beast") is an extinct genus of basal proboscideans from the Eocene of North and West Africa. The first specimen was discovered in strata from the Fayum fossil deposits of Egypt. It was named in 1901 by Charles William Andrews, who suggested that it was an early proboscidean, perhaps ancestral to mastodons, although subsequent workers considered it everything from a relative of manatees to a close relative of both clades' common ancestor. Currently, Moeritherium is seen as a proboscidean that, while fairly basal, diverged before the split between elephantiforms and d
Arsinoitherium
Arsinoitherium (Arsinoe II's beast) is an extinct genus of paenungulate mammals belonging to the extinct order Embrithopoda. It is related to elephants, sirenians, and hyraxes. Arsinoitheres were superficially rhinoceros-like herbivores that lived during the Late Eocene and the Early Oligocene of North Africa from 36 to 30 million years ago, in areas of tropical rainforest and at the margin of mangrove swamps. A species described in 2004, A. giganteum, lived in Ethiopia about 27 million years ago.
Dorudon
Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 41.03 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring long and weighing . Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.
Phiomia
Phiomia (after the Ancient Greek phiom "lake", an ancient name for the Fayum) is an extinct genus of basal elephantiform proboscidean that lived in what is now Northern Africa during the Late Eocene to Early Oligocene some 37–30 million years ago. The type specimen of Phiomia, part of the mandible (lower jaw), was described in 1902 by Charles William Andrews and Hugh John Llewellyn. Unsure of its identity, they assigned it, tentatively, to the obsolete order Creodonta. Subsequently, it was recognised as a proboscidean. Briefly it was treated as a junior synonym of Palaeomastodon, but the two a
Protocetus
Protocetus atavus ("first whale") is an extinct species of primitive cetacean from Egypt. It lived during the middle Eocene period 45 million years ago. The first discovered protocetid, Protocetus atavus was described by based on a cranium and a number of associated vertebrae and ribs found in middle Lutetian Tethyan marine limestone from the Mokattam Formation at Gebel Mokattam near Cairo, Egypt.
Barytherium
Barytherium (meaning "heavy beast") is a genus of an extinct genus of large primitive proboscideans that lived during the late Eocene and early Oligocene in North Africa. The type species is Barytherium grave, found at the beginning of the 20th century in Fayum, Egypt. Since then, more complete specimens have been found at Dor el Talha, Libya. More fossils were also discovered in 2011 in the Aidum area in Dhofar by Oman's Ministry of Heritage and Culture, which was named Barytherium omansi.
Apidium
The genus Apidium (from Latin, a diminutive of the Egyptian bull god, Apis, as the first fossils were thought to be from a type of a cow) is that of at least three extinct primates living in the early Oligocene, from 30 to 28 million years ago. Apidium fossils are common in the Fayoum deposits of Egypt. Fossils of the earlier species, Apidium moustafai, are rare; fossils of the later species Apidium phiomense are fairly common.
Phosphatherium
Phosphatherium escuilliei, named by Gheerbrant, Sudre and Cappetta in 1996, is a basal proboscidean that lived in Africa during the Early Eocene, about 56-55 Ma. It is one of the earliest known proboscideans, together with Eritherium azzouzorum from the Selandian (about 60 Ma). It was found in phosphorites beds from the base of the Ypresian stage of the Ouled Abdoun Basin, which is best known for its exceptionally rich marine vertebrate fauna.
Numidotherium
Numidotherium (from Numidia, and Ancient Greek θηρίον (thēríon), meaning "beast") is an extinct genus of early proboscideans, discovered in 1984, that lived during the middle Eocene of North Africa some 46 million years ago. It was about tall at the shoulder and weighed about .
Titanohyrax
thumb|left|Restoration Titanohyrax is an extinct genus of large to very large hyrax from the Eocene and Oligocene. Specimens have been discovered in modern-day Algeria, Tunisia, Egypt and Libya. Some species, like T. ultimus, are estimated to be as large as the modern rhinoceros. Titanohyrax species are still poorly known due to their rarity in the fossil record.
Daouitherium
thumb|Daouitherium (A) and Numidotherium (B) lower dentition Scale bars: 5 mm
Phiomicetus
Phiomicetus is a genus of protocetid whale that lived between 43 and 42 million years ago during the Lutetian (middle Eocene) period in what is now Egypt. It had powerful jaws and large teeth that would have allowed it to hunt and tear large prey.
Saghacetus
Saghacetus is an extinct genus of basilosaurid early whale, fossils of which have been found in the Upper Eocene (middle Priabonian, ) Qasr el Sagha Formation, Egypt (, paleocoordinates ).
Eosiren
left|thumb|Drawing of skeleton. left|thumb|E. libyca and an extant manatee by Charles R. Knight, 1907 left|thumb|Size of Eosiren (purple) compared to other Eocene sirenians and a human. Eosiren is an extinct genus of sea cow that lived during the Late Eocene (later Priabonian) to Early Oligocene (Rupelian). Several fossils have been found in Egypt. It seems like the species E. abeli were contemporaneous with Protosiren and Eotheroides. Like them, Eosiren closely resembled modern sirenians, with the primary differences being somewhat larger innominates and the presence of thigh bones.
Karanisia
Karanisia is an extinct genus of strepsirrhine primate from Middle Eocene fossil deposits in Egypt.
Apterodon
Apterodon ("without winged tooth") is an extinct genus of hyaenodonts from extinct subfamily Apterodontinae within paraphyletic family Hyainailouridae, that lived from the late Eocene to the early Oligocene epoch in Africa and Europe. Including supplementary materials It is closely related to the African Quasiapterodon.
Stromerius
Stromerius is an extinct genus of basilosaurid early whale known from the Late Eocene (Priabonian, ) of Fayum, Egypt (, paleocoordinates ).
Akhnatenavus
genus of mammals (fossil)
Furodon
Furodon ("tooth of a thief") is an extinct genus of placental mammals from extinct order Hyaenodonta, that lived in North Africa (Tunisia and Algeria) from the early to middle Eocene epoch. It is a monotypic genus that contains the species F. crocheti.
Tutcetus
Tutcetus is an extinct genus of diminutive basilosaurid cetacean from the Bartonian of Egypt. Tutcetus, named after the child pharaoh Tutankhamun, is both one of the oldest known basilosaurids from Africa and the smallest member of the family. It is suggested that the type specimen, a subadult close to maturity, only measured approximately long. The genus is monotypic, only including the species T. rayanensis.
Bothriogenys
Bothriogenys is a genus of anthracotheres that lived in Eastern Africa during the Late Eocene to Early Oligocene.
Masracetus
Masracetus (from Arabic Masr, "Egypt", and Greek ketos, "whale") is an extinct genus of basilosaurid ancient whale known from the Late Eocene (Priabonian, ) of Egypt.
Aegyptocetus
Aegyptocetus is an extinct genus of protocetid archaeocete whale known from Egypt.
Chrysocetus
Chrysocetus is a genus of extinct archaeocete ceteceans belonging to the Basilosauridae that is known from the Late Eocene of the eastern United States and western Africa. Across its range two species are known, the American Chrysocetus healyorum and the African Chrysocetus fouadassii, named in 2001 and 2015 respectively. Chrysocetus was a comparably small basilosaurid, larger than Saghacetus and Tutcetus but smaller than most other members of this group. In addition to being noted for its well-preserved hip bones, Chrysocetus is unique amongst basilosaurids for possibly having only had a sing
Eotheroides
Eotheroides is an extinct genus of Eocene sirenian. It is an early member of the family Dugongidae, which includes the extant dugong. Fossils have been found from Egypt, India, and Madagascar. Eotheroides was first described by Richard Owen in 1875 under the name Eotherium, which was replaced by the current name in 1899.
Afradapis
Afradapis is a genus of adapiform primate that lived during the Late Eocene. The only known species, Afradapis longicristatus, was discovered in the Birket Qarun Formation in northern Egypt in 2009. While its geographic distribution is confined to Afro-Arabia, Afradapis belongs to the predominantly European adapiform family Caenopithecinae. This taxonomic placement is supported by recent phylogenetic analyses that recover a close evolutionary relationship between Afradapis and adapiforms, including Darwinius. While adapiforms have been noted for their strepsirrhine-like morphology, no adapifor
Catopithecus
Catopithecus is an early catarrhine fossil. It is known from more than 16 specimens of a single species, Catopithecus browni, found in the Jebel Qatrani Formation of the Faiyum Governorate, Egypt. The Jebel Qatrani Formation has been divided into two main faunal zones based on the fact that the fauna found in the lower portion of the quarry appear to be more primitive than those found in the upper section. The upper zone has been dated to older than 31 ± 1 myr based on the dating of a basalt layer that lies immediately above the formation and Nicolas Steno's Law of Superposition. The lower zon
Dagbatitherium
Dagbatitherium is an extinct genus of proboscideans. So far a single molar from the phosphate basins of Togo in West Africa has been found. The fossil dates to the Middle Eocene, around 47 million years ago. A striking feature of the tooth are the three pairs of cusps oriented transversely to the longitudinal axis of the tooth. This feature is found in more derived proboscideans, which are grouped in the Elephantiformes. For its age, Dagbatitherium is the earliest elephantiform to date. Furthermore, it is characterized by a low tooth crown and a humped occlusal pattern. The genus was described
Necromantis
Necromantis is an extinct genus of bat that lived during the Eocene. Its fossils are found in the Quercy Phosphorites Formation of France and the Djebel Chambi in Tunisia. Specimens of Necromantis are notable for their large size and specialization towards a predatory lifestyle.
Ancalecetus simonsi
Ancalecetus (from Greek ankale, "bent arm", and ketos, "whale") is an extinct genus of early whale known from the Late Eocene (Priabonian, ) Birket Qarun Formation (, paleocoordinates ) in Wadi Al-Hitan, Egypt. The species is named after anthropologist and primate researcher Elwyn L. Simons who discovered the type specimen in 1985.
Pappocetus
Pappocetus (meaning "grandfather whale") is an extinct protocetid cetacean known from the Eocene of southern Nigeria's Ameki Formation and Togo. More recently, fossil teeth and femurs have also been discovered in the Aridal Formation of the Sahara Desert in southwestern Morocco.
Afrotarsius
Afrotarsius is a primate found in the Paleogene of Africa. left|thumb|Afrasia from Asia and Afrotarsius from Africa exhibit similar morphology of their teeth and lived in the late middle Eocene, suggesting Stem group|stem simians dispersed from Asia to Africa around that time.|alt=Two molars, one of Afrotarsius (left) and one of Afrasia (right), are compared, with an Eocene map of the globe showing where each came from. In the lower left, a life reconstruction of Afrotarsius is shown. The first species to be named, Afrotarsius chatrathi, was named in 1985 on the basis of a single lower jaw fro
Nosmips aenigmaticus
Nosmips aenigmaticus is a rare fossil primate known only from 12 teeth. Most teeth were found at a site in the Fayum Depression about outside Cairo, Egypt.
Anchomomys
Anchomomys is a genus of adapiform primate that lived in Europe and Africa during the middle Eocene.
Boualitomus
Boualitomus ("Bou Ali cutter") is an extinct genus of placental mammals from extinct paraphyletic clade Boualitominae within extinct order Hyaenodonta, that lived in what is now Morocco during the earliest Eocene.
Eocetus
Eocetus (from Ancient Greek ἠώς (ēṓs), meaning "dawn", and Latin cetus, meaning "whale", and thus, "dawn whale") is an extinct protocetid early whale known from the early-late Eocene (Bartonian, ) Giushi Formation in Gebel Mokattam, (, paleocoordinates ) outside Cairo, Egypt. Fossil remains have also been discovered in the Aridal Formation of the Sahara Desert in southwestern Morocco.
Saloumia
Saloumia is an extinct genus of the order Proboscidea. It is one of the oldest members of the order and lived in the middle Eocene of Senegal. It is known only from a single molar, whose pronounced bumpy chewing surface indicates it is probably closely related to Moeritherium.
Algeripithecus
Algeripithecus is an extinct genus of early fossil primate, weighing approximately . Fossils have been found in Algeria and Tunisia dating from 50 to 46 million years ago.
Witwatia
Witwatia (from the Egyptian Arabic Wit Wat meaning "large, flapping wings") is an extinct genus of giant bat that contained two species which lived in the Al Fayyum in Egypt during the late Eocene (Priabonian epoch) and one species which lived in Tunisia during the early Eocene. It is known from a lower jaw and teeth. Three species have been named: the type species W. schlosseri, W. eremicus and W. sigei.
Arcanotherium
Arcanotherium is an extinct genus of early proboscidean belonging to the family Numidotheriidae that lived in North Africa during the late Eocene/early Oligocene interval.
Togocetus
Togocetus ("Togo whale") is a genus of extinct cetacean from the Lutetian (lower Eocene) of Togo, known from a fossilized skeleton discovered a few kilometers north-east of Lomé.
Aegyptonycteris
Aegyptonycteris ("Egyptian bat") is a genus of extinct bat from the Late Eocene of North Africa. It is currently known from a single specimen (holotype CGM 83740) from the Birket Qarun Formation in the Fayum Depression in western Egypt.