Middle Jurassic pterosaurs

Ceoptera (meaning "mist wing") is an extinct genus of darwinopteran pterosaur from the Middle Jurassic Kilmaluag Formation of Scotland. The genus contains a single species, C. evansae, known from a partial skeleton discovered in 2006 and named in 2024. It is the only pterosaur from the Kilmaluag Formation and the second pterosaur named from Scotland, after Dearc in 2022. As one of the only pterosaur skeletons known from the Middle Jurassic, its discovery contributed to understanding the early diversification of the group. Belonging to the group Darwinoptera, a group intermediate between early