Cosmine

Cosmine is a spongy, bony material that makes up the dentine-like layers in the scales of the lobe-finned fishes of the class Sarcopterygii. Fish scales that include layers of cosmine are known as cosmoid scales.

== Description == As traditionally described, cosmine consists of a layer of dentine covered by a continuous sheet of enamel. Pulp cavities, which secrete dentine tubules, are surrounded by a complex polygonal network of 'pore cavities' which pierce the overlying enamel layer, giving cosmine its characteristic dotted appearance. The pulp cavities and pore chambers are connected by a complex, reticulated pore canal network which continues into a layer of vascular bone beneath the dentine. The exact configuration of the pore canal network and shape of the pore chambers differs between various taxa, although the general organization into a single layer of enamel over dentine with pore canals with vascular bone underneath remains consistent, at least within the Sarcopterygii.