trunk

Trunks, also called boles, are the stems of woody plants and the main structural element of trees. The woody part of the trunk consists of dead but structurally significant heartwood and living sapwood, which is used for nutrient storage and transport. Separating the wood from the bark is the cambium, from which trunks grow in diameter. Bark is divided between the living inner bark (the phloem), which transports sugars, and the outer bark, which is a dead protective layer.

The precise cellular makeup of these components differs between non-flowering plants (gymnosperms) and flowering plants (angiosperms). A variety of specialised cells facilitate the storage of carbohydrates, water, minerals, and transport of water, minerals, and hormones around the plant. Growth is achieved by division of these cells. Vertical growth is generated from the apical meristems (stem tips), and horizontal (radial) growth, from the cambium. Growth is controlled by hormones, which send chemical signals for how and when to grow.